Tuzoia

Tuzoia is a large bivalved arthropod having a wide geographic distribution in the strata of the Cambrian Series 2 and 3. One species, Tuzoia bispinosa, is known from abundant material in the Kaili Biota (Cambrian Series 3, Stage 5). Based on 168 specimens from near Balang Village, Jianhe County, Guizhou Province, China, six ontogenetic stages ...

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Tuzoia is a large bivalved arthropod having a wide geographic distribution in the strata of the Cambrian Series 2 and 3. One species, Tuzoia bispinosa, is known from abundant material in the Kaili Biota (Cambrian Series 3, Stage 5). Based on 168 specimens from near Balang Village, Jianhe County, Guizhou Province, China, six ontogenetic stages ...

Pro pokračování aktivujte PREMIUM . Získáte neomezené sledování a stahování 24 hodin denně, 365 dní v roceTuzoia is a relatively common non-biomineralized bivalved arthropod from the Guanshan Biota and, stratigraphically, ranges from Cambrian Series 2 through the Miaolingian Series. Based on new specimens from the Longbaoshan Section of the Wulongqing Formation, this study distinguished and described in detail the Tuzoia in the Guanshan biota. ...Tuzoia is a relatively common non-biomineralized bivalved arthropod from the Guanshan Biota and, stratigraphically, ranges from Cambrian Series 2 through the Miaolingian Series. Based on new specimens from the Longbaoshan Section of the Wulongqing Formation, this study distinguished and described in detail the Tuzoia in the Guanshan biota. ...Jul 28, 2020 - Explore Takashi Ito's board "Cambrian" on Pinterest. See more ideas about prehistoric creatures, prehistoric, prehistoric animals.Soft-bodied fossils of Cambrian age, now known as Burgess Shale–type biotas, were first described from the Parker Slate of the northwest Vermont (USA) slate belt in the late 19 th century, 25 years before the discovery of the Burgess Shale in British Columbia, Canada. Here, we report the rediscovery of fossiliferous horizons at Parker's …Some Kind Of Tuzoia? Some Kind Of Tuzoia? By Nandomas, April 14, 2014 ...对采自云南昆明下寒武统沧浪铺阶乌龙箐组黄色泥岩中的4块Tuzoia标本进行了研究,确定一个新种:Tuzoia kunmingensis sp.nov..新种个体较大,壳面网格清晰,后缘刺保存完好.而其区别于其他种的最主要特征是具5枚基本均匀分布于背缘的背刺.文章简要讨论了Tuzoia的分类位置,认为在软体、附肢还未发现或不十分 ...

The bivalved arthropods Isoxys and Tuzoia with soft‐part preservation from the Lower Cambrian Emu Bay Shale Lagerstätte (Kangaroo Island, Australia) - …Tuzoia is a large bivalved arthropod having a wide geographic distribution in the strata of the Cambrian Series 2 and 3. One species, Tuzoia bispinosa, is known from abundant material in the Kaili Biota (Cambrian Series 3, Stage 5). Based on 168 specimens from near Balang Village, Jianhe County, Guizhou Province, China, six ontogenetic stages ...A fragment of a Burgess Shale-type fossil, bivalved arthropod Tuzoia Walcott, 1912, is described from shales of the mid-Cambrian Buchava Formation in the Skryje-Týřovice Basin, Central Bohemia, Czech Republic. It is the second recorded find of Tuzoia from West Gondwana. Uncrushed and uncoloured preservation of the tuzoiid valve is ...首次报道山东潍坊馒头组上部(寒武系第三统)的东北吐卓虫Tuzoia manchuriensis,该种的模式标本产自辽宁省烟台镇东南3.2km的当十岭,但保存不很完整。潍坊标本在后腹缘处有两个明显、粗壮的边缘刺,前腹缘处有许多呈锯齿状小刺,这是辽宁标本所没有保存的特征,是对模式种的重要补充。Tuzoia is a large, bivalved Cambrian arthropod having a widespread distribution in shallow‐shelf to deeper marine Lagerstätten; stratigraphically it ranges from Cambrian Series 2 through the ...Tuzoia is retrieved by a Bayesian phylogenetic analysis as an early mandibulate hymenocarine lineage, exemplifying the rapid diversification of this group in open marine environments during the ...Tuzoia is a relatively common non-biomineralized bivalved arthropod from the Guanshan Biota and, stratigraphically, ranges from Cambrian Series 2 through the Miaolingian Series.

3.. Materials and methodsTwo specimens of Tuzoia and two specimens of anomalocaridid appendages housed in the University of Kansas Museum of Invertebrate Paleontology, Biodiversity Research Center (KUMIP 298507a, 298527a, 307020b, 298529a) from the Early and Middle Cambrian were selected for the taphonomic investigation. A specimen of Canadaspis (KUMIP 307021a) from the Middle Cambrian horizon ...Beside the Tuzoia, Anomalocaris, and merostome described from this formation in this paper several other unusual fossils have been found, including notably a sponge and a species of some Holothurian possibly belonging to the genus Peytonia. Both the eastern and western Mesonacid faunas have now furnished such fossils that clearly show that the ...Tuzoia is retrieved by a Bayesian phylogenetic analysis as an early mandibulate hymenocarine lineage, exemplifying the rapid diversification of this group in open marine environments during the ...Tuzoia was a large bivalved arthropod witch existed widely in the Cambrian seas all over the world. Two species of Tuzoia from Lower Cambrian Guanshan Fauna of Eastern Yunnan were systematically studied in this thesis, including one species Tuzoia sinensis redescribed and the other species indeterminate Tuzoia sp. described and compared.The reticulated surface on the large carapaces of Carnarvonia (pl. 33, fig. 1) and Tuzoia (pl. 33, fig. 2), also approximates in pattern and size that of Nebaliopsis typica Sars. The merostomes Sidneyia, Amiella, Habelia (pl. 29), Molaria (pl. 29), and Emeraldella (pl. 30), have a compact cephalic shield and well-defined thoracic and abdominal …

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Tuzoia is a relatively common non-biomineralized bivalved arthropod from the Guanshan Biota and, stratigraphically, ranges from Cambrian Series 2 through the Miaolingian Series. Based on new ...New Data on the Bivalved Arthropod Tuzoia From the Cambrian (Series 2, Stage 4) Guanshan Biota in Kunming, Yunnan, Southwest China Published in: Frontiers in Earth Science, April 2022 DOI: 10.3389/feart.2022.862679: Authors: Yichen Wu, Jianni Liu View on publisher site Alert me about new mentions.Tuzoia, and several species of isoxyids and hymenocarines (electronic supplementary material, Analyses 2.2). The same analysis was run using an alternative dataset with a more conservative coding ...The results of the more conservative dataset (dataset 2) are shown as an inset containing only the closest allied taxa to Tuzoia (see electronic supplementary material, figure S6 for full tree).

Tuzoia is a relatively common non-biomineralized bivalved arthropod from the Guanshan Biota and, stratigraphically, ranges from Cambrian Series 2 through the Miaolingian Series. Based on new ...Taxonomy from Vannier et al. (2007, p. 458). Vannier et al. 2006 assume that Tuzoia and the also Cambrian genus Isoxys are possibly representatives of the class Thylacocephala. Revised diagnosis for the genus Tuzoia Walcott, 1912 in Vannier et al. (2007, pp. 458-459): "Large bivalved arthropod (v...Sep 1, 2014 · 3.Soft anatomy of Isoxys minor. Isoxys minor is the most commonly represented species in the Guanshan fauna, which is widely distributed in the Wulongqing Formation (Cambrian Series 2, Stage 4; Palaeolenus Zoon) at the Shitangshan Section near Gaoloufang and Gangtoucun villages, Kunming City, Lihuazhuang, Yiliang County, Shijiangjun and Sapushan, Wuding County (Luo et al., 2008). Tuzoia retifera Walcott = Tuzoia retifera (Walcott, 1912) . 1912 69 Walcott Cambrian Geology and Paleontology II plate 33 - P77YRK from Alamy's library of millions of high resolution stock photos, illustrations and vectors. Shop Alamy Prints Store - Crafted by Experts, Delivered to Your Door.11 thg 1, 2020 ... Francis Mavinga Tuzola. 1988 - 2019. Photos. Obituary ... Click here to send a condolence to the family of Francis Tuzola, or to share a memory.This is a rare arthropod, apparently bivalve, known as Tuzoia that is found from several of the world's Cambrian Lagerstätten, such as Canada's Burgess Shale (Tuzoia burgessensis, Tuzoia retifera and Tuzoia canadensis) and China's Maotianshan Shales (Tuzoia sinensis), and still others from the Eager Formation in Canada and Emu Bay Shale in Australia.Tuzoia are interpreted as nektonic animals that probably inhabited the lower level of the water column in shallow shelf settingsatdepthsofperhaps100-150morless ...ãeeãeeãeeãee All items are 100% Authentic and accurately described!! Always looking for other collectors to buy our mineral, and receive your positive feedback in repeat, We will do anything we can doCopeland, M.J. 1993: Contributions to Canadian Paleontology Anomalocaris and Tuzoia from the lower Cambrian Eager Formation near Cranbrook, British Columbia Geological Survey of Canada Bulletin 1993(444): 1-5The morphology of Tuzoia is reinterpreted in the light of abundant new specimens from the Middle Cambrian Burgess Shale (British Columbia, Canada) and Kaili (Guizhou, China) Lagerstätten. Tuzoia was a very large (up to 180 mm long) bivalved arthropod with a nonmineralized domelike carapace strengthened by prominent pointed features and often flanked by a lateral ridge bearing a spiny frill ...

Media in category "Tuzoia" The following 7 files are in this category, out of 7 total. Artistic reconstruction of the Cambrian (Drumian) ...

Tuzoia, and several species of isoxyids and hymenocarines (electronic supplementary material, Analyses 2.2). The same analysis was run using an alternative …Tuzoia retifera (Q2461984) From Wikidata. Jump to navigation Jump to search. No description defined. edit. Language Label Description Also known as; English: Tuzoia retifera. No description defined. Statements. instance of. taxon. 0 references. image. Tuzoia retifera USNM PAL 57720.jpg. 0 references. taxon name.Tuzoia represents the largest bivalved arthropod from the Spence Shale, with some valves reaching 12 cm long by 8 cm wide. Radiodonts are also fairly common in the Spence Shale and at least three species are known, an indeterminate Anomalocaris species ( Briggs et al . 2008 ), Caryosyntrips camurus ( Pates & Daley 2017 ) and at least one ...Tuzoia is a relatively common non-biomineralized bivalved arthropod from the Guanshan Biota and, stratigraphically, ranges from Cambrian Series 2 through the Miaolingian Series. Based on new ...Tuzoia – a rare and bizarre Arthropod from the Cambrian of China. Beautiful result after a challenging transfer-preparation. Note the reddish color!Tuzoia is a large, bivalved Cambrian arthropod having a widespread distribution in shallow‐shelf to deeper marine Lagerstätten; stratigraphically it ranges from Cambrian Series 2 through the ...Tuzoia australis, the palaeoscolecid priapulid Palaeoscolex antiquus, and two animals of uncertain affinities, Myosco-lex ateles and Vetustovermis planus. The report of Anomal-ocaris from the Emu Bay Shale (McHenry and Yates 1993) was followed by the recognition of two species in that genus (Nedin 1995a), one of which was formallyThe Guanshan Biota (Cambrian Series 2, Stage 4) is a typical Burgess Shale-type biota and is one of the most significant Cambrian Konservat-Lagerstätten from China. Tuzoia is a relatively common ...

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Nov 13, 2009 · The eyes in Tuzoia sp. were more than twice as large to those in Tuzoia australis and show evidence of a visual cortex, but were located in a similar position, at least in dorsoventral specimens (EXPLANATION OF PLATE 4 , EXPLANATION OF PLATE 6 ). Tuzoia is a relatively common non-biomineralized bivalved arthropod from the Guanshan Biota and, stratigraphically, ranges from Cambrian Series 2 through the …Billy Tuzola Dunga Daniel is on Facebook. Join Facebook to connect with Billy Tuzola Dunga Daniel and others you may know. Facebook gives people the...Fibulacaris nereidis is unique in its carapace morphology and overall widens the ecological disparity of Cambrian arthropods and suggests that the evolution of a 'bivalved' carapACE and an upside-down lifestyle may have occurred early in stem-group crustaceans. The origin of the arthropod carapace, an enlargement of cephalic tergites, can be traced back to the Cambrian period. However, its ...This widespread distribution resembles that of nektonic Peytoia, which occurs across Laurentia (Daley et al., 2013a; see also Pates et al., 2018), South China (Liu, 2013 our unpublished data) and Baltica (Daley and Legg, 2015) and, to some extent, that of the presumed pelagic Isoxys and Tuzoia (Vannier and Chen, 2000; Vannier et al., 2007).Tuzoia getzi Resser, 1929 - fossil bivalved arthropod from the Cambrian of Pennsylvania, USA. (YPM 10044, Peabody Museum of Natural History, Yale University, New Haven, Connecticut, USA) (centimeter scale) This fossil is from the Kinzers Lagerstätte, a soft-bodied fossil deposit in the Lower Cambrian of Pennsylvania. About 60 species are present in the interval that contains exceptional ...Vannier et al. 2006 assume that Tuzoia and the also Cambrian genus Isoxys are possibly representatives of the class Thylacocephala. Emmended diagnosis for the genus Isoxys by Garcia-Bellido 2009, p. 1224: "Arthropod with one pair of cephalic appendages and a uniform series of at least 13 pairs of biramous appendages. Long, narrow body covered ...Tuzoia was first described by Charles Walcott (1912) based on a single carapace specimen from the Burgess Shale. He collected many more specimens after this publication that were later described in detail by Resser (1929). Resser designated ten different species of Tuzoia, including specimens from the Burgess Shale, the Eager Formation of ...S u nZ X,W a n gP La n dY u a nJ L(2015) The first complete Tuzoia manchur- iensis from the Cambrian series 3 of Weifang, Shandong Province. Acta Palaeontologica Sinica 54 ,1 1 3 – 19 (in ... ….

Merkmale. Tuzoia retifera hatte einen eiförmigen Umriss (Verhältnis Länge zu Höhe etwa 1,45) und eine Panzerlänge von 20 bis 120 mm. Der obere Rand war gerade bis leicht konvex und hatte keine Stacheln oder Dornen. Das vordere Rostrum war breit angelegt, sehr kurz, nicht spitz und leicht nach unten gerichtet. Die Sehkerbe war gut ausgebildet. Das hintere Rostrum war sehr klein und stumpf.Subtaxa: Tuzoia australis. View classification. Ecology: nektobenthic omnivore. Fossilworks hosts query, analysis, and download functions used to access large paleontological data sets. It presents taxonomic, distributional, and ecological data about the entire fossil record. Gateway to the Paleobiology Database ...Check out my latest presentation built on emaze.com, where anyone can create & share professional presentations, websites and photo albums in minutes.The association of trace fossils and non-biomineralized carapaces has been reported from Cambrian Lagerstätten worldwide, but the abundance, ichnodiversity, taphonomy and ecological significance of such associations have yet to be fully investigated. Two main end-member hypotheses are explored based …Tuzoia parva (Walcott); Briggs 1977:618, pl. 72, fig. 8, text-fig. 25; Lieberman 2003:678; Vanier et al. 2007:467. Catalog record: USNM PAL 57716 Original historic description: PAGE Hymenocaris parva Walcott 185 Fig. 7. (×4.) Type specimen of the species and genus, showing several appendages and the abdomen - P77FBP from Alamy's library of ...Nov 1, 2009 · Abstract: Abundant material from a new quarry excavated in the lower Cambrian Emu Bay Shale (Kangaroo Island, South Australia) and, particularly, the preservation of soft‐bodied features previously unknown from this Burgess Shale‐type locality, permit the revision of two bivalved arthropod taxa described in the late 1970s, Isoxys communis and Tuzoia australis. The collections have also ... This specimenbearing interval, about one meter thick, also yields abundant articulated trilobites, the large bivalved arthropod Tuzoia manchuriensis (Sun et al., 2015), threedimensionally ...Tuzoia was a diverse genus widely distributed across Laurentia, Bohemia,Siberia,andGondwana.Withatleast14validspecies,it ranks among the most diverse non-biomineralized or lightly Tuzoia, Wang et al. (2010) described a new species of Isoxys, I. shandongensis, from the upper part of the Manto Formation at Linyi, Shandong Province (Figs 1, 2). This locality also yielded abundant ..., The origin of mandibulates, the hyperdiverse arthropod group that includes pancrustaceans and myriapods, dates back to the Cambrian. Bivalved arthropod groups such as hymenocarines have been argued to be early mandibulates, but many species are still poorly known, and their affinities remain uncertain. One of the most common and globally distributed Cambrian bivalved arthropods is Tuzoia ..., Tuzoia - from Mount Tuzo, in the Valley of the Ten Peaks, named in 1907 after Henrietta Tuzo, who was the first to climb this mountain.. burgessensis - from Mount Burgess (2,599 m), a mountain peak in Yoho National Park. Mount Burgess. The name was given in 1886 by Otto Klotz, the Dominion topographical surveyor, after Alexander Burgess, a former Deputy Minister of the Department of the ..., PubMed, Tuzoiids like Tuzoia burgessensis here would have grown up to about 23cm long (~9″). They had large eyes on short stalks, a pair of simple antennae, a horizontal fluke-like tail fan, and twelve pairs of appendages along their body - with the front two pairs at the head end being significantly spinier, and most (or all) of these limbs also ..., fossil huntress musings meant to captivate, educate and inspire, Tuzoia was gifted a redesign after various studies, link in description. 9. 1 commento. Condividi. Salva. 7. Postato da 10 mesi fa., Jul 14, 2015 · The morphology of Tuzoia is reinterpreted in the light of abundant new specimens from the Middle Cambrian Burgess Shale (British Columbia, Canada) and Kaili (Guizhou, China) Lagerstätten. Tuzoia was a very large (up to 180 mm long) bivalved arthropod with a nonmineralized domelike carapace strengthened by prominent pointed features and often ... , It is thought that Tuzoia species probably lived in the water column, perhaps explaining their great distributions. Note: Marks on all scale bars are centimeters (cm). Tuzoia guntheri Robison and Richards, 1981. KUMIP 134797 (Paratype; part, counterpart). Marjum Formation, House Range, Millard County, Utah. Figure 8.4 in Robison & Richards (1981)., Tuzoia is a relatively common non-biomineralized bivalved arthropod from the Guanshan Biota and, stratigraphically, ranges from Cambrian Series 2 through the Miaolingian Series. Based on new specimens from the Longbaoshan Section of the Wulongqing Formation, this study distinguished and described in detail the Tuzoia in the Guanshan Biota., Fibulacaris nereidis is unique in its carapace morphology and overall widens the ecological disparity of Cambrian arthropods and suggests that the evolution of a 'bivalved' carapACE and an upside-down lifestyle may have occurred early in stem-group crustaceans. The origin of the arthropod carapace, an enlargement of cephalic tergites, can be traced back to the Cambrian period. However, its ..., Vannier et al. 2006 assume that Tuzoia and the also Cambrian genus Isoxys are possibly representatives of the class Thylacocephala. Emmended diagnosis for the genus Isoxys by Garcia-Bellido 2009, p. 1224: "Arthropod with one pair of cephalic appendages and a uniform series of at least 13 pairs of biramous appendages. Long, narrow body covered ..., palaeothalassia and the first Wuliuan occurrence of Tuzoia guntheri. We also report on a new hurdiid carapace element and additional specimens of Buccaspinea cooperi?, Dioxycaris argenta, Hurdia sp., and Tuzoia retifera. This new material improves our understanding of the panarthropod fauna of the Spence Shale Lagerstätte and substantially ..., Tuzoia is a large, bivalved Cambrian arthropod having a widespread distribution in shallow‐shelf to deeper marine Lagerstätten; stratigraphically it ranges from Cambrian Series 2 through the ..., 112 Tuzoia canadensis 2. 113 Tuzoia? parva 2. 114 Tuzoia praemorsa 2. 115 Tuzoia retifera 2. 116 Waptia fieldensis 2. 117 Yohoia tenuis 2. 118 Chancia palliseri 2. 119 Ehmaniella burgessensis 2 ..., Tuzoia polleni. Geological Time: Early Cambrian. Size (25.4mm=1 inch): 74 mm long by 45 mm across on an 150 mm by 138 mm matrix. Fossil Site: Eager Formation, Cranbrook, British Columbia, Canada. Description: This unusual fossil is an example of a rarely seen Phyllocarid known as Tuzoia polleni. ..., Tuzoia is retrieved by a Bayesian phylogenetic analysis as an early mandibulate hymenocarine lineage, exemplifying the rapid diversification of this group in open marine environments during the ..., Media in category "Tuzoia" The following 7 files are in this category, out of 7 total. Artistic reconstruction of the Cambrian (Drumian) ..., Tuzoia australis, the palaeoscolecid priapulid Palaeoscolex antiquus, and two animals of uncertain affinities, Myosco-lex ateles and Vetustovermis planus. The report of Anomal-ocaris from the Emu Bay Shale (McHenry and Yates 1993) was followed by the recognition of two species in that genus (Nedin 1995a), one of which was formally , Tuzoia is a relatively common non-biomineralized bivalved arthropod from the Guanshan Biota and, stratigraphically, ranges from Cambrian Series 2 through the Miaolingian Series. Based on new ..., Tuzoia is a large, bivalved Cambrian arthropod having a widespread distribution in shallow-shelf to deeper marine Lagerstätten; stratigraphically it ranges from Cambrian Series 2 through the Miaolingian Series. In the present study, three species are recognized from relatively deep-water deposits of the Balang Formation (Cambrian Stage …, Redlichiais a genus ofredlichiidtrilobitein the familyRedlichiidae, with large to very large species (up to 35 centimetres or 14 inches long). Fossils of various species are found in Lower Cambrian (T, The bivalved arthropods Isoxys and Tuzoia (Lower and Middle Cambrian) are two other potential thylacocephalan candidates making this group of arthropods a ..., A fragment of a Burgess Shale-type fossil, bivalved arthropod Tuzoia Walcott, 1912, is described from shales of the mid-Cambrian Buchava Formation in the Skryje-Týřovice Basin, Central Bohemia, Czech Republic. It is the second recorded find of Tuzoia from West Gondwana.Uncrushed and uncoloured preservation of the tuzoiid valve is consistent with the very thin non-mineralised cuticle ..., Copeland, M.J. 1993: Contributions to Canadian Paleontology Anomalocaris and Tuzoia from the lower Cambrian Eager Formation near Cranbrook, British Columbia Geological Survey of Canada Bulletin 1993(444): 1-5 , Isoxys (meaning "equal surfaces") is a genus of extinct bivalved Cambrian arthropod; the various species of which are thought to have been freely swimming predators. It had a pair of large spherical eyes (which are the most commonly preserved feature of the soft-bodied anatomy), and two large frontal appendages used to grasp prey., The largest Emu Bay Shale Tuzoia specimens with circular eyes preserved in situ show that these visual organs do not reach more than 9 mm in diameter (21, 26), and even the biggest Tuzoia ..., Tuzoiids like Tuzoia burgessensis here would have grown up to about 23cm long (~9"). They had large eyes on short stalks, a pair of simple antennae, a horizontal fluke-like tail fan, and twelve pairs of appendages along their body - with the front two pairs at the head end being significantly spinier, and most (or all) of these limbs also ..., Beside the Tuzoia, Anomalocaris, and merostome described from this formation in this paper several other unusual fossils have been found, including notably a sponge and a species of some Holothurian possibly belonging to the genus Peytonia. Both the eastern and western Mesonacid faunas have now furnished such fossils that clearly show that the ..., Tuzoia is a large bivalved arthropod having a wide geographic distribution in the strata of the Cambrian Series 2 and 3. One species, Tuzoia bispinosa, is known from abundant material in the Kaili Biota (Cambrian Series 3, Stage 5)., A new leanchoiliid is described, Yawunik kootenayi gen. nov, based on 42 specimens from the newly discovered Marble Canyon locality of the Burgess Shale, which stresses the historical importance of polarization over data content in scenarios of early arthropod evolution and a pressing need to investigate the impacts of coding inapplicables, especially given the inflating effect of implied weights., 东北吐卓虫(Tuzoia manchuriensis)完整个体在山东潍坊寒武系的首次发现. 首次报道山东潍坊馒头组上部(寒武系第三统)的东北吐卓虫Tuzoia manchuriensis,该种的模式标本产自辽宁省烟台镇东南3.2km的当十岭,但保存不很完整。. 潍坊标本在后腹缘处有两个明显、粗壮的 ... , The arthropods belong to Isoxys mackenziensis n. sp., Tuzoia cf. T. guntheri Robison and Richards, 1981, Branchiocaris? sp., Perspicaris? dilatus Robison and Richards, 1981, and bradoriids, along with fragments of arthropods of indeterminate affinities. The style of preservation indicates that most soft parts underwent complete biodegradation ...